Brochis Sub Clade 4 is by far the largest of the sub clades with 32 species and twice as many coded species. These are what are comonly known as straight snouted species and some of the largest species to be found within Corydoradinae.
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(Steindachner, 1877)
(Sands, 1995)
(Cope, 1872)
(Nijssen & Isbrücker, 1983)
(Elwin,1939)
Bentley, Grant & Tencatt, 2021
(Nijssen, 1972)
Tencatt & Ohara, 2016
(Nijssen & Isbrücker, 1980)
(Grant, 1998)
Sands, 1995
(Nijssen & Isbrücker,, 1983)
Meinken, 1957
Meinken, 1957
(Nijssen, 1972)
(Castro, 1986)
(Knaack, 1962)
Nijssen, 1970
(Nijssen & Isbrücker, 1983)
(Burgess, 1993)
(Knaack, 2004)
(Nijssen & Isbrücker, 1986)
(Myers, 1933)
(Knaack, 2004)
(Nijssen & Isbrücker, 1976)
(Weitzman & Nijssen, 1970)
(Dinkelmeyer, 1995)
Isbrücker & Nijssen, 1973
Burgess, 1983
Nijssen & Isbrücker, 1980
Dinkelmeyer, 1996
Knaack, 1999
Weitzman, 1960
Burgess, 1993
Sands, 1995